The department offers a one-quarter course (Neurobiology 206) on the structure and function of the nervous system, which is open to medical and graduate students and advanced undergraduates.

Advanced courses are open to students who have completed the basic course.
To sign up for courses, go to the Axess website.


NBIO 101/201. Social and Ethical Issues in the Neurosciences---Influences on public debate and policy of scientific advances in the study of the brain and behavior: theories of brain function; philosophical and scientific approaches; advances in the neurosciences, possible uses in medical therapy, and interventions involving genetic screening, genetic selection, enhancement of neurological functioning, and manipulation of behavior; questions related to medical therapy, social policy, and broader considerations of human nature such as consciousness, free will, personal identity, and moral responsibility. May be taken for 2 units without a research paper. Prerequisite: Neuroscience, Biology, or Symbolic Systems major; or Human Biology core; or consent of instructor.   

2-4 units, Spr (Hurlbut, W)


NBIO 198. Directed Reading in Neurobiology---Prerequisite: consent of instructor. (Staff)

1-18 Units, Aut, Win, Spr, Sum (Staff)


NBIO 199. Undergraduate Research---Investigations sponsored by individual faculty members. Prerequisite: consent of instructor.

1-18 Units, Aut, Win, Spr, Sum (Staff)


NBIO 206. The Nervous System---Introduction to the structure and func­tion of the nervous system, including neuroanatomy, neurophysiology, and systems neurobiology. Topics include the properties of neurons and the mechanisms and organization underlying higher functions. Framework for general work in neurology, neuropathology, clinical medicine, and for more advanced work in neurobiology. Lecture and lab components must be taken together.

7-8 units, Win (Raymond, J)


NBIO 206A. The Nervous System: Lab---Enrollment limited to Neurosciences graduate students. Lab component of NBIO 206; MD students enroll in NBIO 206. Provides a survey of human neuroanatomy, with a focus on gross anatomy and histology. Students work with human brain samples. Open to Neuroscience Program graduate students only.

3 Units, Win (Raymond, J)


NBIO 216. Genetic Analysis of Behavior---(Same as MCP 216.) Advanced seminar. Findings and implications of behavioral genetics as applied to invertebrate and vertebrate model systems. Topics include biological clocks, and sensation and central pattern generators. Relevant genetic techniques and historical perspective. Student presentation.

4 units, not given this year


NBIO 218. Neural Basis of Behavior---Advanced seminar. The principles of information processing in the vertebrate central nervous system, and the relationship of functional properties of neural systems with perception and behavior. Emphasis is on the visual and auditory systems. Original papers; student presentations. Prerequisite: 206 or consent of instructor.

4 units, alternate years, not given this year


NBIO 220. Central Mechanisms in Vision-based Cognition---Contem­porary visual neuroscience, emphasizing the neural mechanisms underly­ing primate vision and visually guided behavior. Seven foundational topics in visual neuroscience; current papers concerning each topic. Student presentations. Computer-based demonstration exercises.

2-4 units, Aut (Moore, T; Newsome, W)


NBIO 227. Understanding Techniques in Neuroscience---This student-organized and student-taught course examines and discusses a wide spec­trum of methods commonly used in the field of neuroscience. Techniques discussed range from molecular/genetic to electrophysiology and whole brain imaging. Prepares students to understand common techniques used in any neuroscience paper, seminar, or poster. Each class includes an over­view lecture, brief presentations by senior graduate students who practice these techniques, and relevant examples from the literature. Optional laboratory demonstrations outside of class are provided.

2 units, Aut (Newsome, W)


NBIO 228. Mathematical Tools for Neuroscience---Student-instructed. For students with no math background beyond basic calculus, or as a review for more advanced students. Techniques useful for analysis of neural data including linear algebra, Fourier transforms, probability and statistics, signal detection, Bayesian inference, and information theory.

2 units, Spr (Baccus, S)


NBIO 254. Molecular and Cellular Neurobiology---(Same as BIOSCI 154/254.) For advanced undergraduates and graduate students. Cellular and molecular mechanisms in the organization and functions of the nervous system. Topics: wiring of the neuronal circuit, synapse structure and synaptic transmission, signal transduction in the nervous system, sensory systems, molecular basis of behavior including learning and memory, molecular pathogenesis of neurological diseases. Prerequisite for undergraduates: Biological Sciences core or equivalent, or consent of instructors.

4-5 units, Aut (Luo, L; Shen, K; Clandinin, T), alternate years, not given next year


NBIO 258. Information and Signaling Mechanisms in Neurons and Circuits---(Same as MCP 258.) How synapses, cells, and neural circuits process information relevant to a behaving organism. How phenomena of information processing emerge at several levels of complexity in the nervous system, including sensory transduction in molecular cascades, in­formation transmission through axons and synapses, plasticity and feedback in recurrent circuits, and encoding of sensory stimuli in neural circuits.

4 units, Aut (Baccus, S)


NBIO 299. Directed Reading in Neurobiology---Prerequisite: consent of instructor.

1-18 units, Aut, Win, Spr, Sum (Staff)


NBIO 370. Medical Scholars Research---Provides an opportunity for student and faculty interaction, as well as academic credit and financial support, to medical students who undertake original research. Enrollment is limited to students with approved projects.

4-18 Units, Aut, Win, Spr, Sum (Staff)


NBIO 399. Graduate Research---Allows for qualified students to un­dertake research sponsored by individual faculty members. Prerequisite: consent of instructor.

1-18 units, Aut, Win, Spr, Sum (Staff)


See respective department listings for course descriptions. See degree requirements above or the program’s student services office for applicabil­ity of these courses to a major or minor program.


MCP 215. Synaptic Transmission,

5 units, Aut (Staff)


PSYCH 204A. Computational Neuroimaging

1-3 units, Spr (Wandell, B)


PSYCH 250. High-level Vision,

1-3 units, alternate years, not given this year